Biodiversity & Landscape Modeling Group

Journal Publications

	67.	He, F. Maximum entropy, logistic regression, and species abundance. Oikos (accepted).
	66.	Gray, L. and He, F. Spatial point-pattern analysis for detecting density-dependent competition in a boreal chronosequence of Alberta. Forest Ecology and Management (in press).
	65.	Zillio, T. and He, F. Inferring species abundance distribution across spatial scales. Oikos (in press).
	64.	Yavitt, J.B., Harms, K.E., Garcia, M.N., Wright, S.J., He, F. and Mirabello, S.J. 2009. Spatial heterogeneity of soil chemical properties in a lowland tropical moist forest, Panama. Australian Journal of Soil Research 47:674-687.
	63.	Morlon, H., White, E.P., Etienne, R.S., Green, J.L., Ostling, A., Alonso, D., Enquist, B.J., He, F. , Hurlbert, A., Magurran, A.E., Maurer, B.A., McGill, B.J., Olff, H., Storch, D. and Zillio, T. 2009. Taking species abundance distributions beyond individuals. Ecology Letters 12:488-501.
	62.	Hu, X.-S., He, F. and Hubbell, S.P. Communtiy differentiation on landscapes: drift, migration, and speciation. Oikos 118:1515-1523 (pdf).
	61.	He, F. 2009. Price of prosperity: economic development and biological conservation in China. Journal of Applied Ecology 46:511-515 (pdf).
	60.	Babak, P. and He, F. 2009. A neutral model of edge effects. Theoretical Population Biology 75:76-83 (pdf).
	59.	Legendre, P., Mi, X.C., Ren, H.B., Ma, K.P., Sun, I.F., Yu, M.J. and He, F. 2009. Partitioning beta diversity in a subtropical broad-leaved forest of China. Ecology 90:663-674 (pdf).
	58.	Lin, K., Zhang, D.-Y. and He, F. 2009. Demographic trade-offs in a neutral model explain death rate-abundance rank relationship. Ecology 90:31-38 (pdf).
	57.	Chen, X.-Y. and He, F. 2009. Speciation and endemism under the model of island biogeography. Ecology 90:39-45 (pdf).
	56.	Li, L., Huang, Z.-L., Ye, W.-H., Cao, H.-L., Wei, S.-G., Wang, Z.-G., Lian, J.-Y., Sun, I.-F., Ma, K.-P. and He, F. 2009. Spatial patterns of tree species in a subtropical forest of China. Oikos 118:495-502 (pdf).
	55.	Hubbell, S.P., He, F., Condit, R., Borda-de-Água, L., Kellner, J. and ter Steege, H. 2008. How many tree species are there in the Amazon and how many of them will go extint? PNAS 105:11498-11504 (pdf).
	54.	Babak, P. and He, F. 2008. Species abundance distribution and dynamics in two locally coupled communities. Journal of Theoretical Biology 253:739-748 (pdf).
	53.	Gamarra, J.G.P. and He, F. 2008. Spatial scaling of mountain pine beetle infestations. Journal of Animal Ecology 77:796-801 (pdf) (Aber News;BBC news)
	52.	Zhu, H.T., He, F. and Zhou, J. 2008. Auto-multicategorical regression model for the distribution of vegetation. Statistics and Its Interface (pdf).
	51.	Getzin, S., Wiegand, T., Wiegand, K. and He, F. 2008. Heterogeneity influences spatial patterns and demographics in forest stands. Journal of Ecology 96:807-820 (pdf)
	50.	Shen, T.-J. and He, F. 2008. An incidence-based richness estimator for quadrats sampled without replacement. Ecology 89:2052-2060 (pdf).
	49.	Witte, J.-P. M., He, F. and Groen, C. 2008. Grid origin affects scaling of species across spatial scales. Global Ecology and Biogeography 17:448-456 (pdf).
	48.	He, F. and Tang, D.L. 2008. Estimating the niche preemption parameter of the geometric series. Acta Oecologica 33:105-107 (pdf).
	47.	Nelson, W., Potapov, A., Lewis, M., Hundsdorfer, A. and He, F. 2008. The balance of complexity in mechanistic modeling: Risk analysis in the mountain pine beetle. Journal of Applied Ecology 45:248-257 (pdf).
	46.	Pueyo, S., He, F. and Zillio, T. 2007. The maximum entropy formalism and the idiosyncratic theory of biodiversity. Ecology Letters 10:1017-1028 (pdf).
	45.	Hu, X.-S., Yeh, F.C. and He, F. 2007. Sex-ratio distortion driven by migration loads. Theoretical Population Biology 72:547-559 (pdf).
	44.	Hu, X.-S., He, F. and Hubbell, S.P. 2007. Species diversity in local neutral communities. American Naturalist 170:844-853 (pdf).
	43.	He, F. and Gaston, K. 2007. Estimating abundance from occurrence: an underdetermined problem. American Naturalist 170:655-659 (pdf).
	42.	McGill, B.J., Etienne, R.S., Gray, J.S., Alonso, D., Anderson , M.J., Benecha, H.K., Dornelas, M., Enquist, B.J., Green, J., He, F., Hurlbert, A., Magurran, A.E., Marquet, P.A., Maurer, B.A., Ostling, A., Soykan, C.U., Ugland, K., White, E. 2007. Species abundance distributions: moving beyond single prediction theories to integration within an ecological framework. Ecology Letters 10:995-1015 (pdf).
	41.	Getzin, S., Dean, C., He, F., Trofymow, T., Wiegand, K. and Wiegand, T. 2006. Spatial patterns and competition of tree species in a Douglas-fir chronosequence on Vancouver Island. Ecography 29:671-682 (pdf).
	40.	Hu, X.-S., He, F. and Hubbell, S.P. 2006. Neutral theory in population genetics and macroecology. Oikos 113:548-556 (pdf).
	39.	Magnussen, S., Pélissier, R., He, F. and Ramesh, B.R. 2006. An assessment of sample-based estimators of tree species richness in two wet tropical forest compartments in Panama and India. International Forestry Review 8:417-431 (pdf).
	38.	Gaston, K.J., Borges, P.A.V., He, F. and Gaspar, C. 2006. Abundance, spatial variance, and occupancy: species distribution in the Azores. Journal of Animal Ecology 75:646-656 (pdf) .
	37.	Wills, C., Harms, K.E., Condit, R., King, D., Thompson, J., He, F., Muller-Landau, H.C., Ashton, P., Losos, L., Comita, L., Hubbell, S.P., LaFrankie, J., Bunyavejchewin, P., Dattaraja, H.S., Davies, S., Esufali, S., Foster, R., Gunatilleke, N., Gunatilleke, S., Hall, P., Itoh, A., John, R., Kiratiprayoon, S., Loo de Lao, S., Massa, M., Nath, C., Noor, S., Kassim, A.R., Sukumar, R., Suresh, H.C., Sun, I.-F., Tan, S., Yamakura, T. and Zimmerman, J. 2006. Non-random processes contribute to the maintenance of diversity in tropical forests. Science 311:527-531 (pdf).
	36.	Hu, X.-S. and He, F. 2006. Seed and pollen flow in expanding a species' range. Journal of Theoretical Biology 240:662-672 (pdf).
	35.	Volkov, I., Banavar, J.R., He, F., Hubbell, S.P. and Maritan, A. 2006. Comparing models of species abundance: a reply. Nature 441:E1-E2 (pdf).
	34.	Muller-Landau, H.C., Condit, R.S., Harms, K.E., Marks, C.O., Thomas , S.C. , Bunyavejchewin, S., Chuyong, G., Co, L., Davies. S., Foster, R., Gunatilleke, S., Gunatilleke, N., Hart, T., Hubbell, S.P., Itoh, A., Kassim, A.R., Kenfack, D., LaFrankie, J.V., Lagunzad, D., Lee, H.S., Losos, E., Makana, J.-R., Ohkubo, T., Samper, C., Sukumar, R., Sun, I.-F., Nur Supardi M.N., Tan, S., Thomas, D., Thompson, J., Valencia, R., Vallejo, M.I., Muñoz, G.V., Yamakura, T. Zimmerman, J.K., Dattaraja, H.S., Esufali, S., Hall, P., He, F., Hernandez, C., Kiratiprayoon, S., Suresh, H.S., Wills, C. & Ashton, P. 2006. Comparing tropical forest tree size distributions with the predictions of metabolic ecology and equilibrium models. Ecology Letters 9:589-602.
	33.	He, F. and Hubbell, S.P. 2005. Percolation theory for the distribution and abundance of species: a reply. Physical Review Letters 95, article 189804 (pdf).
	32.	Volkov, I., Banavar, J.R., He, F., Hubbell, S.P. and Maritan, A. 2005. Density dependence explains tree species abundance and diversity in tropical forests. Nature 438:658-661 (pdf).
	31.	He, F. and Hu, X.-S. 2005. Hubbell's fundamental biodiversity parameter and the Simpson diversity index. Ecology Letters 8:386-390 (pdf).
	30.	Hu, X.-S. and He, F. 2005. Background selection and population differentiation. Journal of Theoretical Biology 235:207-219 (pdf).
	29.	He, F. 2005. Deriving a neutral model of species abundance from fundamental mechanisms of population dynamics. Functional Ecology 19:187-193 (pdf)
	28.	He, F., Gaston, K.J., Connor, E.F. and Srivastava, D. 2005. The local-regional relationship: immigration, extinction, and scale. Ecology 86:360-365 (pdf).
	27.	Wu. J., Huang, J.-H., Han, X.-G., Gao, X.-M., He, F., Jiang, M.-X., Jiang, Z.-G., Primack, R. B. and Shen, Z.-H. 2004. Three Georges Dam: An ecological perspective. Frontiers in Ecology and the Environment 2:241-248 (pdf).
	26.	He, F. and Hubbell, S.P. 2003. Percolation theory for the distribution and abundance of species. Physical Review Letters 91, article 198103 (pdf).
	25.	He, F. and Gaston, K.J. 2003. Occupancy, spatial variance, and the abundance of species. American Naturalist 162:366-375 (pdf).
	24.	Trofymow, J.A., Addison, J., Blackwell, B.A., He, F., Preston, C.A. and Marshall, V.G. 2003. Attributes and indicators of old-growth and successional Douglas-fir forests on Vancouver Island. Environmental Reviews 11:S187-S204.
	23.	He, F., Zhou, J. and Zhu, H.T. 2003. Autologistic regression model for the distribution of vegetation. Journal of Agricultural, Biological and Environmental Statistics 8:205-222 (pdf).
	22.	He, F., LaFrankie, J.V. and Song, B. 2002. Scale dependence of tree abundance and richness in a tropical rain forest, Malaysia. Landscape Ecology 17:559-568 (pdf).
	21.	Gaston, K.J. and He, F. 2002. The distribution of species range size: a stochastic process. Proceedings of the Royal Society of London, B 269:1079-1086 (pdf).
	20.	He, F. and Legendre, P. 2002. Species diversity patterns derived from species-area models. Ecology 83:1185-1198 (pdf).
	19.	He, F., Gaston, K.J. and Wu, J. 2002. On species occupancy-abundance models. ÉcoScience 9:119-126 (pdf).
	18.	Holt, A.R., Gaston, K.J. and He, F. 2002. Occupancy-abundance relationships and spatial distribution: a review. Basic & Applied Ecology 3:1-13.
	17.	He, F. and Gaston, K.J. 2000. Estimating species abundance from occurrence. American Naturalist 156:553-559 (pdf).
	16.	He, F. and Duncan, R.P. 2000. Density-dependent effects on tree survival in an old-growth Douglas-fir forest. Journal of Ecology 88:676-688 (pdf).
	15.	He, F. and Alfaro , R.I. 2000. White pine weevil attack on white spruce: a survival time analysis. Ecological Applications 10:35-42 (pdf).
	14.	He, F. and Gaston, K.J. 2000. Occupancy-abundance relationships and sampling scale. Ecography 23:503-511 (pdf).
	13.	He, F. and Barclay, H. 2000. Long-term response of understory plant species to thinning and fertilization in a Douglas-fir plantation on southern Vancouver Island , British Columbia. Canadian Journal of Forest Research 30:566-572 (pdf).
	12.	Tomlin, E.S., Alfaro , R.I., Borden, J.H. and He, F. 1998. Histological response of resistant and susceptible white spruce to simulated white pine weevil damage. Tree Physiology 18:21-28.
	11.	He, F., Legendre, P. and LaFrankie, J.V. 1997. Distribution patterns of tree species in a Malaysian tropical rain forest. Journal of Vegetation Science 8:105-114 (pdf).
	10.	Alfaro, R.I., He, F., Tomlin, E. and Kiss, G. 1997. White spruce resistance to white pine weevil related to bark resin canal density. Canadian Journal of Botany 75:568-573.
	9.	He, F. and Alfaro, R.I. 1997. White pine weevil (Coleoptera: Curculionidae) attack on white spruce: spatial and temporal patterns. Environmental Entomology 26:888-895 (pdf).
	8.	He, F. and Legendre, P. 1996. On species-area relations. American Naturalist 148:719-737 (pdf).
	7.	He, F., Legendre, P. and LaFrankie, J.V. 1996. Spatial patterning of diversity in a tropical rain forest in Malaysia. Journal of Biogeography 23:57-74 (pdf).
	6.	Alfaro, R.I., He, F., Kiss, G., King, J. and Yanchuk, A. 1996. Resistance of white spruce to white pine weevil: Development of a resistance index. Forest Ecology and Management 81:51-62.
	5.	Jeglum, J. and He, F. 1995. Pattern and vegetation-environment relationships in a boreal forested wetland in northeastern Ontario. Canadian Journal of Botany73:629-637.
	4.	He, F., Legendre, P., Belleheumer, C. and LaFrankie, J.V. 1994. Spatial scale and diversity patterns: a study of a tropical rain forest of Malaysia. Environmental and Ecological Statistics 1:265-286.

Book Chapters

	3.	He, F. and Condit, R. 2007. The distribution of species: occupancy, scale and rarity. Pages 32-50 in Scaling Biodiversity, eds. D. Storch, P.L. Marquet and J.H. Brown. Cambridge University Press (pdf).
	2.	Borda de Agua, L., Hubbell, S.P. and He, F. 2007. Scaling biodiversity under neutrality. Pages 347-375 in Scaling Biodiversity, eds. D. Storch, P.L. Marquet and J.H. Brown. Cambridge University Press (pdf).
	1.	He, F. and Reed, W. 2006. Downscaling abundance from the distribution of species: occupancy theory and applications. Pages 89-108 in Scaling and Uncertainty Analysis in Ecology: Methods and Applications, eds. J. Wu, B. Jones, H. Li and O. Loucks. Springer, New York (pdf).